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Acyl-CoA synthetase long-chain family member 4 (ACSL4) catalyzes the conjugation of PUFAs to coenzyme A, forming PUFA-CoAs, which are subsequently incorporated into membrane phospholipids by lysophosphatidylcholine acyltransferase 3 (LPCAT3) ( Lipid metabolism interacts with ferroptosis through multiple pathways: Stearoyl-CoA desaturase-1 (SCD1) generates monounsaturated fatty acids that substitute for PUFAs, thereby reducing lipid peroxidation substrates